Zookeepers have paid emotional tribute to the world’s oldest known Sumatran orangutan, “a grand old lady” who died at a Western Australian zoo on Monday. Puan was given to Perth zoo in 1968 and is believed to have been born in Sumatra 1956. At 62 years she lived well beyond her typical life expectancy and was recognised by the Guinness Book of Records as the oldest verified member of her species in 2016.
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